Botrychium dissectum  (B. biternatum; B. oneidense; Sceptridium dissectum) cut-leaf grape fern Ophioglossaceae BODI; NY; Q, j; R, gb;

 

Botrychium dissectum.D. Busemeyer.SEINet.swbiodiversity.org

Botrychium dissectum.D. Busemeyer.SEINet.swbiodiversity.org (Accessed 8/2015).

Botrychium dissectum is a fern 10-30 cm tall, from fleshy, mycorrhizal roots, it appears to be colonial (personal observation). 

Leaf solitary, arising from lower part of fertile stem, roughly triangular in outline, on a stalk 3-7 cm long, blade divided three to four times, often reddish when expanding, and bronze in autumn, divisions usually unequal, irregular, tips of major divisions less divided; fertile spike (sporophyll) to 10 cm long, fertile frond branched and much divided. 

Spore cases yellow-brown; Aug-Nov. 

Wetland status: FAC. 

Frequency in NYC: Very infrequent. 

Origin: Native. 

Habitat: Open areas or woods on sterile soil. 

Notes: Gametophyte non-photosynthetic, closely associated with specific fungi below ground similar to Lycopodium sp. (S. Clemants, personal communication; Montgomery and Fairbrothers 1992). It has been shown that, for some species of Botrychium, exotic earthworms may inhibit establishment of new individuals (Gundale 2002). This includes the Botrychium/Sceptridium dissectum var. obliquum found by Zihao Wang in Manhattan.

Botrychium dissectum.© Lawrence S. Gilliam 2009. hosted by the USDA-NRCS PLANTS Database

Botrychium dissectum.© Lawrence S. Gilliam 2009. hosted by the USDA-NRCS PLANTS Database